A Hitherto的意思|示意

In 51 survivors and their descendants, they discovered a hitherto-unknown variant of PRNP, the gene which makes prions, the proteins that spread the disease.

他们在51位幸存者和他们的后代身上发现了一种目前为止无人发现的PRNP变体。PRNP是含有库鲁病传染源——朊病毒——一种蛋白质的源代码。

A hitherto polite dispute is acquiring a sharper edge.

迄今为止貌似斯文的争端正在变得尖锐有棱。

As a result, a myriad of hitherto separate objects are becoming connected to networks, from televisions and cars to industrial machinery and farmland.

结果，无数个迄今为止还是相互分离的事物被网络连接起来，从电视、汽车到工业机械和农田。

On April 5th it emerged it had detected what looked like a hitherto unknown particle.

4月5日暴露出它已探测到一个好像迄今未知的粒子。

The gland has a hitherto unknown structural organization.

腺体有一个前所未有的组织结构。

In April a hitherto obscure Icelandic mountain called Katla leapt from nowhere to favourite, with odds of 7/4, following the eruption of its previously less-active neighbour, Eyjafjallajokull.

四月份冰岛的一个名叫Katla的鲜为人知的火山从无人问津一跃成为热门，赔率在7/4，一般会在与其毗邻的火山Eyjafjallajokull爆发之后喷发。